flujo de energia amb terres y acuatic

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  • 8/6/2019 Flujo de Energia Amb Terres y Acuatic

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    Energy flow within terrestrial and

    aquatic ecosystems Energy available for growth and reproduction (as a

    function of food availability):

    = GPE - NPE

    Tells usamount of energy flow

    Not rate at which energy flows

    Ecosystem and community development is

    regulated by therate of energy flow as well

    How fast does available energy flow between

    trophic levels

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    Second question of energy flow within a system is: how fast does energy transfer occur?

    Residence time = inverse of energy transfer rate RT = biomass / net productivity

    RT (yr) = energy stored in biomass (kJ m )-2

    Net productivity (kJ m yr )-2 -1

    Measure of biomass accumulation within any given trophic level

    (g(C) m )

    (g(C) m yr )

    -2

    -2 -1or

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    Wet tropical rainforest:

    Produce about 1800 g m-2 yr-1

    andLiving average biomass of 42 000 g m-2

    42 000 g m-2

    1800 g m-2 yr-1= RT = 23 yr

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    Temperate scrubland:

    Produce about 600 g m-2 yr-1

    and

    Living average biomass of 6 000 g m-2

    6 000 g m-2

    600 g m-2

    yr-1

    = RT = 10 yr

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    Open ocean:

    Produce about 125 g m-2 yr-1

    andLiving average biomass of 3 g m-2

    3 g m-2

    125 g m-2 yr-1= RT = 0.02 yr

    (About 9 days)

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    Ricklefs (1972)

    Peat bogs near Root Springs, Concord MA

    Herbivores: assimilate energy at a rate of 0.31 J m-2 s-1

    Aquatic plants and algae produce 0.09 J m-2 s-1

    So, where is the other 0.22 J m-2 s-1 coming from?

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    Ecosystems do not exist

    individually autochothonous production = energy fixed

    within the system

    allochothonous production = energy fixed

    outside of the system

    energy can be from input from external

    system

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    General Trends

    assimilation efficiency increases at higher trophiclevels

    NPE decreases at higher trophic levels

    GPE decreases at higher trophic levels

    Ecological efficiency averages about only 10%

    GPE(n) / GPE (n-1) = efficiency of transfer

    between specific trophic levels

    Aquatic systems typically support more trophic

    levels than terrestrial systems

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    Nutrient cycles

    Biogeochemical sources

    Biotic = nutrients released from decomposers

    Abiotic = nutrients released from erosion of

    substrate or fixed by primary producers Indirect = nutrients stored in soil

    Direct = nutrients absorbed directly from

    decomposers Indirect = higher nutrient capital

    Amount of stored nutrients within the system

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    Atmosphere

    Primary

    producers

    ConsumersDead organic

    material

    Soil

    Indirect nutrient cycling

    Nutrients can flow from source to source

    Length of time at each source dependant

    upon source characteristics

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    Atmosphere

    Primary

    producers

    ConsumersDead organic

    material

    Soil

    Direct nutrient cycling

    Nutrient uptake increased by presence

    of mycorrhizae (fungal producers)