AUdlondl:Tu1nl:fliluli ob/k &d6 Technical Paper no. 16/2006
Inland Fisheries Research and Development Bureau
Department of Fisheries
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EFFECT OF FEEDING FREQUENCY ON GROWTH PERFORMANCE OF
coMMoN LOWLAND FROG, Rana rugulosa (wiegmann) TADPOLE
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Anuwat Uppanunchai
Ratana InsiripongTheerachai Pongjanyakul
Ministry of Agriculture and Cooperatives
ส้านัก
วิจัยและพัฒ
นาประมงน้้าจืด
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Phrae Inland Fisheries Research and
Development Center
Amphoe Nongmuangkhai, Phrae 54170
Tel.0-5463-5024
2006
Technical Paper no. 16/2006
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COMMON LOWLAND FROG, Rana rugulosa (Wiegmann) TADPOLE
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ส้านัก
วิจัยและพัฒ
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EFFECT OF FEEDING FREQUENCY ON GROWTH PERFORMANCE OF
coMMoN LOWLAND FROG, Rana rugulosa (wiegmann) TADPOLE
Anuwat Uppanunchai
Phrae Inland Fisheries Research and Development Center,
Amphoe Nongmuangkhai, Phrae 54170, Thailand
Ratana InsiriPong
Khon Kaen Inland Fisheries Research and Development Center,
Amphoe Mueang, Khon Kaen 40000, Thailand
Theerachai Pongj anYakul
ubonratchathani Inland Fisheries Research and Development center,
Amphoe Mueang, Ubonratchathani 34000, Thailand
ABSTRACT
Nursing of common lowland frog with five feeding frequencies 2,3, 4, 5 and 6
times/day was studied by using the average initial weight of tadpole 0'24+0'01 g' The
experiment was conducted in 45x90x45 cm glass aquaria, contained 50 litres of water at
stocking density of50 tadpoles/aquarium and fed to satiation at each feeding frequency for 4
weeks.
The results showed that feeding frequency had strongly effect on survival rate.
The survival rate of frog fed at the rate of 5 and 6 times/day were significantly higher than
the frog fed 2,3 and 4 times/day (p<0.05). Thus the feeding frequency at least 5 times/day is
suggested for nursing offrog (tadpole).
Key words: tadpole Rana rugulosa feedtng frequency
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l.3o+o.o6o"
1 .1 4t0. 1 2"
1.26*0.o3"o
0.23*o.oro
o.2o*o.o l"o
o.zo*o.o2"o
0. I 9+0.00'
0.20+0.02"
8.oo*0.580
5. I 5t0.25"
3.90+0. I 9b
3.35*0.33n
2.64+0.31"
58.00+2.00"
72.67*t.15'
86.67+8.33'
97.33+3.06u
98.67*2.3ru
z
3
4
5
6
z.i4+o.roo
2.55+0.04'
3.01+0.07"
2.95+0.10"
3.42+o.1ro
0.24*o.o l o
0.23+0.01"
0.24*o.o l"n
0.23+0.01'
0.26+0.01"
4.44+0.1 8'
3.05+o.l ro
1.9710.06"
1.56*0.o6o
1.25+0.04"
47.33!3.06^
58.6714. I 6b
62.67+3.060
86.67+2.31"
88.0015.29'
z
3
4
5
6
2.39+0. I 9"n
2.43+0.06t
2.25+0.38'b
z.o2+0.13^
2. I 4+0.06"b
0.1 5+0.01"
0.1 5*o.ooo"
0.1 5*o.o 1o'
0. I 3+0.00'
0.14*o.o l t
3.21+0.240
2.06+0.05"
t.66+0.25b
1.29+0.08"
L 1 2+0.05'
30.00+2.00'
39.33+3.06'o
35.33*l.r5o
54.0014.00'
63.3314. I 6'
2
3
4
J
6
4.52+9.331
4.25+0.06'b
3.67+0.t 1"
3.72+O.lg^
3.96*0.o9"n
0.21+o.o4u
0.19+0.00"b
0. I 7+0.00"
0. I 710.00"
0. 18+0.01"'
2.28+0.03"
r.5z*o.o2u
1. I 4+0.01 "
0.88*o.03n
0.76+0.03"
25.33+1. I 5"
26.67+2.31'
30.67+l . I 5"
47.33+6.430
54.67*7.02'
flilrutfi9t, i i. - , u v dt a
ri t tq a t.tr n r n u o x uo n u : n 1 u 1 0 { n q u m a l{ n u n 1 il [[u
I a ooirrn nsiri riuvrr.: ao fr fr :v 6'u fiudr ri'iy o.os
.r 1 u ii'r.l o r #rG I r riu ttafl {'i r fi nr r r ruส้านัก
วิจัยและพัฒ
นาประมงน้้าจืด
8
L v ier d o u d'r u n r t ru 6 lu n r : 1 #o r u r : si r I ri'u s : v 6'u rfl u
:euut?al + tt'rjqr#
ay . a .uu nr:"[r.f orvl: (n5{/au)
! --A tt Arirr rin sr":rEu 6u rqd u (nfu ) 0.24+0.01 0.24+0.01
4.25+0.06" 3.67+0.11^
0.19+0.00^ 0.17+0.00"
0.06+0.00b 0.04+0.00"
1 .4t 1.09
0.24+0.01 0.24+0.01
3.72+O.lg" 3.96+0.09b
0.17+0.00" 0.18+0.01"
0.03+0.00" 0.03+0.00"
0.81 0.76
! -- s) Au"rruna?d9rvt lu[aa u (nflJ)
'
,t , , - t d
il:Ntolo'1ilt:tl tl{9tost ?glo?u
(nfrul6'rliu)iq , r '
rl?ursuo r14r: fr "[fi siooi'rsionfI 0.10+0.02"
(nfirtnf.:)
d'qr:rnr:'l#orur:r 4d , ! u u il
( ul o f rfi u pilri rm fin 6'rrn :-.r )
6'q:r:oq (%)
0.24+0.01
5.00+0.01d
o.2l+0.04^
2.00
25.33+1.15^ 26.61+2.31^ 30.67+1.15^ 47.33+6.43b 54.67+7.02b
t i i. - Y u u 4t u I d t o aa
?m.nutfiE ritrafiufirirfi'tJ6'rud'nu:nruro'rnqufrnt{nuflltJttu?uouttdo.l'l'liln'l1tJttflnfl1{numl{dCIqli-Yl:v9tiluud]nflJ 0.05
z. rJ?r.noro rn1: dl#Ausio fra sioiu
z. t rl?:L rnr o r u r : fi l#n^u si o si'r si o{u
rl?rurororur:fi1#fiusio6"lsio{u{osnuurfioqrradxun?rild"lunr:'l#orur: 2, 3, 4, s
I - d! v | / d ra jq gl+ I 9-,!- y-- -^y-^- v t l4 t
rray 6 n:{/'lu nlu:yuu[']ar + #:Jqril fi:J?rurororu1:vrtrnunosi'lqioSu rd'lmnoo.l t ii'rlnrri frrit
rvirri':r 0.23+0.01,0.20+0.01,0.20+0.02,0.19+0.00 [[au 0.20+0.02 nfil srrrLaird'u (srr:rld r) tdo
s) t ) y iq ct, I .- d t^ jqga11Odoilll',r.ldilSt-trd',t ilu'jlnuulfrouura6?un'rllJolunl: [f orur: 2 n:{/?u lJ1l:tllil0'11'{r:vltl'lflu
qio6'rsioiururnn'jrodr.:firiudrni'cymr.rafin'1p<0.05) riunrurfioUUrad?unxrilfi 5 rrav o n9.rl5u rrqi
rrflneir{rj'uodr':kifiriuiird'omrqafisr^ (p>0.05) nuurdouuradlun'lrild 3 LLav 4n9.rriu ,Jodnr"'lfi
ruav 6 o#oliu fir]?rLrarorrr:fi1#fiusio6'rsioiufiorrru
r r fl n ei r.1 ri'u o d r I hi fi riu d I d'q lt t s a o^ or^ (p> 0 . 0 5 )
r,rd'ruqao{ z dil.lltyi rl?rurnrorrr:fi1#fiuqiod'rqioiufirirtvitril o.z++0.01, 0.23+0.01,
0.24+0.01, 0.23+0.01 rray 0.26+0.01 nfrJ qrrudrd'rL (nr:tfi t) tttiomqaolmrlan^6tt#': uu'jrnuurfiy iq q9 I .- 4 la ---+q y^ I u .r-!'-.- ---^ I 4
ouutadxuq?1ilil'[unt:'[riorur: 6 ni{/iu ilt-Jiruratotrl:m [f nun0nx0t0']uillnm(fl 0u1.1u
,iua',d'rur',uafio^ 1p<o.os) d.rdnuurfiorluradron?1ildlunr:1#orur: 2, 3 trau + ng.rlfu traunuu'l
A gt j6r qil - J - d la jq/^ I u drlouilrao?un'nrJ6rtunr:1#orr.rr: 3, 4 t[au s nYltiu firl?rurarorur:fr'lr{ilusroql"]eioiuilnrru
", h 14 q
r ra n ei r { riu o d r.r't ri fi riu d r ri'q vr r'r a fi 6 (p>0. 0 5 )
ส้านัก
วิจัยและพัฒ
นาประมงน้้าจืด
9
ua'{mqao{ s xi'ilor# rl?rursuorr.rr:fi1#fiusio6"rsioiufirirrvirfi'u o.tsto.ol, 0.15+0.00,
0.r5+0.0r, 0.13+0.00 rlou 0.14+0.0t nfru srrruiirdr (r',r',od t) u.laomqaouvrr.rafio^rrdr vr:L'irnuurfi
ouurad'runrrrL6lunr:1#ormr: s u9otiu firJ:^rurorerr,rr:fil#fiusio6':sioiurloufi4o otjxfi
;;;ril;;,;;;;.. ,s; vT.rdnuurfiorpradroo.',ud'lun',r1#orur: 2,3 $av+ u#.,riu ttasnilulA gt jq ql J .- d la jqga r-ir'- | u dmouuraq?unxrilo tunr:1#orur: 3, 4 rray o nfrtiu frrl?rLriuorur:fr"[rinusiosr":sioiurLnrtru
" , o r h td v
r r qr n si r r ri'u o d r {'[ il fi fi u d t ri'iy m t t a o^ ot^ ( p > 0. 0 5 )
ud'{rqao{ + &ilet'ty" rl?ruruorrr:fi1#fiusio6":qioiufirirtvirri'u o.zt+0.04. 0.19+0.00.
0.17+0.00, 0.17+0.00 rlas 0.18+0.01 nfru srrruiird'u (qrr:xfi t) riliovterdoumrlafisi^rrdr mu'jrnuurd
ourradrun:rrufi1unr:1#orur: z.9rl5, firlirurororur:fil#fiusie6'rsioiutrnn'jroeixfiriudrd'q
rNdfia (p<0.05) rlunlurfiorlurafirunrrild + ,,u, s orlliu ttsittsrnsirlri'uodr.rkifirludrri'q1vrr'l
afior^ 1p>0.05) nuurfiorlurad'runrrild : ,,u, o ogoliu ,iudnrulfiuquradrtlnrrrufilunr:l#u.r*',,,
3, 4, srray 6 nfrl5u firl?rLrarorur:fi1#fiusio6"lsio{ufinrru,,rnrir,,i'uotjrlhifiritldrriqlvrr.rafio
b>0.0s)z.z 6'fl:rnr:1#orur:
6fl:rnr:'l#orilr: tul ofr6urilrirurin6"rlof.r) nuunru-,durllrndrunrrru6lunr:1#3 - dt at r 4-
1#orilt:ud'.:vrerao{lii'ilert#firiro-tl.t'l: 2,3, 4,5 ttau 6 nt{/iu tilu:gugt:a1 4 dil91114 ilOA:1fl1:r ld d o
rvirri':t s.00t0.58, 5.15+0.25, 3.90+0.19, 3.35+0.33 LLav 2.64+0.:: uJoftfiuri srrrLdrdl (nr:rtfi t)i aa c., , j iq qY - !.- du qe)
rfromnaouvn{dddrldx iluitnuulyroUUran?1iln lunt: lfi01r1: 2 n:{/?u lJ0n:1n1: tllo'l?11:j'
urnmqqrrflnnr{our{rlriudrri'rymxafior^ (p<0.05) :oraruliln^onru",duqr",udrunrrru6lunr:1#! - 3: A v ru61unr:1#orilr:4tmvso9otirfidn:rnr:1#otur:fi011{1: 3 n:{/XU ll{UnUU]llOUUlAqXUnXl
axrilrrflnsil,rfiuodr{lxifiriotirri'eyvrr.rafio^ 1p>0.05) rtrunuurfiorlura6'ruurlud'lunr:'[#orr.rr: o
u#utiu fi6'qr:rnr:1#orrr:rloufiqorrnnoit.rotir.:firiudrri'cg1l1{dfiei'$<0.05)
ild'wqao{ z &ilq:rii 6'sr:rnr:1#orrr:firirrvirri'r 4.44+0.18, 3.05+0.11, 1.97+0.06,
t d d d o ' -oo 9t t
1.56+0.06 lrau 1.25+0.04 lilo:rryun fl]il6itetu (srr:xfi t) rfromeraotvtr{afiet^rrdr urL'jrnuurfiouula
u.',ud1rn',r1#orr.rr: z ufotiu fi6'n:rnr:l#u",r',ru',ndqorusrnsitodtrfirierdrri'iyvrr{dil^et^ (p<0.05)
h ra A I io qe) J - e Y:o.ra.r'lr]fionuurfroqrtadrunrrru6'lunt:1#orut: 3.4, 5 ttav e oflliu qrtuiitli'u
?id'{roao{ : ti'rlflr# d'sr:rnr:1#orur:firirrvirrir 3.21+0.24, 2.06+0.0s, 1.66+0.2s,
t d d d o ' oo / ' '
1.29+0.08 rray 1.12+0.05 [il0:rsiua fltilareiu (flr:r{fi 1) tnoovrodolvrr.:an^fittdt vlu'irnuurfioUUtai. qs) J - dv qY j I
n?tilolunt:1r.Torur: z nflriu ru0fl:rnr: tilor14'1:rJlnvrqnouxfiriudrri'qmrlao'o^ 6<0.05) ,u.,uo
'lrjn^onuurfiourrad':unrrrLfilunr:1#orur: 3 rtav + o#oriu mrruiird'u cirunuurdorlurad:a
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atr^or^ 1p>o.os;
ส้านัก
วิจัยและพัฒ
นาประมงน้้าจืด
10
ud'{rqao{ + d'rlqr# dsr:rnr:1#orur:finirrvirriu 2.28+0.03, t.5z+o.oz, 1.14t0.01,
0.88+0.03 rrau 0.76+0.0r ulofr6ud nrruii16'r (nr:r.rd t) ,fiu*raurmr.rafisr'rtda vlu'jrnuurdoqilloi" 6il J - dv qs) i
n?1il6i tun1: tHor1.i1: 2 nf{/iu ildrl:rnr:-[rolilrTilrnmqqrtflnnr{odr.rfiriodrriqvrf i?ffiet^ ,r.O.O!h fd A y j6l q, - !, .- o u
:o{a{ tilnonilulvrou1jlafl?un?llJo lunl: l#otvtl: 3,4,5 $aY o nfl/iu qrrrua{rdu
3. fln:t:ooA e.) iq qY J - dr -r 4
fllJU1llOUIJtaq',tufl'JtrJ0 tUnl:1#01il 11 2,3,4, 5 rrAv O n:-lriU tilU:suvtrar + d:lflrfdurloet:t:oerfia{mooo.r, Alr'.lri,rirri'l st.oo+2.00,72.67+1.15,86.67+8.33,97.33+3.06lrax98.67+2.31
r rd d o v i i .aa il . ' i ' fru6'[Un1i1#rrloirfrur,l atUiitftU (ort:tlvt l) tilovl9ldoU1ll.:d0fltta2 ?'llJ?lnUu11101-{Ulaq?Un?1
or1.r1: z n9.riiu fid'fl:r:oeeirfido rloan'jrodr'rfiriotirri'q1vrxafin^ (p<0.05) rirLnuurfior.lulad'lu
n?ril6 3, 4, 5 rrav 6 ng{/{u,,ur,fiuurndu noo nuu'rfiouurad'lunrrildlunr:"1#orurt 3 trae 49'!u a I 4 il Aq :1#orrr:5rrauonirliufid'sr:r:oofin?rilltflnsir{fi'unr{/?u d?unlju11]ouulao?un?]il0 tun'l
o tir.rhifi riu dr6'q1 m r r an^fi tr>O. 05 )@t 4u
ua{ilqao{ z&ilafi6'sr:r:oqfirirrvirri':r 47.33+3.06,58.67!4.16,62.6i+3.06,86.67+2.31
ruay 88.00+5.29 ulofrduoi snileird'u (srr:r.rd t) rdovreaouvrrlan^sr^ttd? v{u'itnuu',durlurodrtlio qe) J - du i i , I 1.- 4v--.e,-9^.--^ -332
" ' '
n?1il0 tunri tilor4r: z nfrriu fro'er:r:oerrirfiqo rioan'jrotjrrfiriodtri'rgmxafin^ (p<0.05) fr"rfi
- -d '--^-.r'.ru61unr:1#orur: : LLay 4 u9uliu fidqr:r:onfinx1ilttnnei1.r6'uodr.r'hifinilu111 0 1.{ U 1A q',r Ufi',l
riudrri'rgmr.rd6r^er^ (p>0.05) rravfioqura6'runrrrudlunr:1#ornr: s rtav o ofoliu ildsr:r:onfinrril
r r fl n ei 1 { ri'u o d r.r hj fi rT0 ?i r ri'q m r'r a n^ fi (p> 0 . 0 5 )
ud'rmoaor a ai'rlor# dfl:r:oflfirirrvirri'u 30.00+2.00, 39.33+3.06, 35.33+1.15, 54.0014.00
tlay 63.33!4.16 riloirdud'nril6ii6'il (rlr:rqd t) urioueaoumr{afl^erttd'r vlu'irnuurdorlurnd'lui" qe I - du i
nxrilrrtunr: tlrorrr: 5 [rau 6 nf{/iu r]on:r:onrulnrqq rutnn'jtodrrfirludrri'qvn'ido^sr^ (p<o os)I ! ) , iq qv 3 .- 4u d I n rdvr'lfrnuurfroUU1n6?un'lrrLil"[unr:]r{or?n: 2 LLay 3 n5{/au trofl:r:0fliln'ltilttanfll{nu0u'l.1 [lJtJ
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r rn n ei r { rTu o d r .r hi fi riu d r 6ry m r.r a fi o^ b>0. 0 5 )
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LLay 54.6i+7.02 ulefr6up{ qrrrLd16'r (srr:rrfi z) rruiouodouur{afifirr#r uu'jrnuurfiouuradlg
n'nildlunr:1#orrr: 5 rrav 6 ugoliu fio'sr:r:oo'hirrnnsirrri'uodr.:firiodrflcyvn.rdo^er^ 1pro.osl ,,rifiI
d'm:r:oournn'irnrurdouurad?un'lru61unr:1#or?n: 2,3 LLae + nf':tiu oeir.:firiudrri'qmr'idfiet'e ' i e/ i" q9 3.- dv d
(p<0.05) vi.rfinuurfioU:l'lafr?on'lrru6lunr:l#or?n: 2, 3 tLa-- + nfqliu ilo'rr:rnr::oqfrn:ril
r rn n ei 1 { ri'u o ei r .: hi fi riu d r ri'iy vr r'r a fi 6 (p>0. 0 5 )
ส้านัก
วิจัยและพัฒ
นาประมงน้้าจืด
ll
94o4. nildiluglu1
Y g) f,q q, 3 - 4qnraurin^rir:ufi'i1{nt:oUUtanu6?un?rilir.."[un1:]fiormr: 2,3,4,5 ttav 6 nt{/iu tiluu t J t o 3 o , 'cl t -- ^ - d 4 d I d
:uuyr?ar + drJqril vlu'jrqnru4il^uo{ilrfid1oqjludrr 26.0-28.4 o{fllrryorfrud nxliltilun:otilueil{il
rirotjlurir.r 7.59-8.01 rJ?ruro{oonGroufiavarorfrfirirodlurir{ 4.0-5.s ru^aa^n:-ru/fisr:g'
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ส้านัก
วิจัยและพัฒ
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ส้านัก
วิจัยและพัฒ
นาประมงน้้าจืด
13
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rugul osa,Wiegmann). no lrl: vru lri'.1dr, nturl:uil{. n?{tlluq. t :o u #r'u t 4 .a r u e y- - 4 !r' --i a ' ltUf. 2539.ntfl{ [fl?faurutfrqa, qdun uqticy, nrru ariatu'1olt, ?:u ']ry:n: tulu ltav u?dilil ?'1.1f
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I a I -- Iil5ulJ.t, lJu t-11u4 utnu gl: fl l?{91:. nE {[vlv'|"l. 255 YIu1.
Boujard. T.lgg9. Feeding behavior and regulation of food intake. 1n Nutrition and
Feeding of Fish and Crustacean. edited by Guillaume J., S. Kaushik, P. Bergot and
R. Metailler. Praxis Publishing Ltd, UK. pp.17-25.
BrettJ. R. and T. T. D. Groves. t979.Digestion. In FishPhysiologyvol. VII. Physiological
Energetics. Edited by Hoar W. S., D. J. Randall and J. R. Brett. Academic Press, Inc.
pp.280-344.
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